Ecosystem
An ecosystem is all the living organisms in an area together with the nonliving parts of their environment, linked by flows of energy and matter.
Definition
An ecosystem is a place where living things and nonliving things work together, including the plants, animals, and microbes plus the sunlight, water, soil, and air they depend on, and it can be as small as a puddle or as large as the whole ocean as long as its parts stay connected. More formally, an ecosystem is a community of organisms (biotic factors) interacting with each other and with nonliving abiotic factors such as sunlight, temperature, water, and soil, through which energy flows one way, from the sun through producers to consumers and decomposers, while matter such as carbon, nitrogen, and water cycles within it. At the functional level, an ecosystem is characterized by energy flow, biogeochemical cycling, trophic structure, and emergent properties such as productivity, resilience, and nutrient retention, where gross primary production is the total energy fixed by autotrophs and net primary production, what remains after their own respiration, sets the energy budget for every trophic level above; ecosystems are open systems that exchange matter and energy with their neighbors and respond to disturbance through succession, and the ecosystem services they provide, pollination, water purification, carbon storage, climate regulation, underlie much of human economic activity.
Example
A pond ecosystem has fish, frogs, insects, algae, and bacteria living together with the water, mud, sunlight, and rocks, and in a North Carolina hardwood forest oaks and hickories capture sunlight, squirrels and deer eat acorns and leaves, owls and foxes eat the squirrels, and fungi and bacteria return nutrients from fallen leaves to the soil, all sustained by roughly $110$ to $130\ \text{cm}$ of rain per year. Net primary production ranges from about $2{,}200\ \text{g}$ of dry biomass per square meter per year in tropical rainforest to under $100$ in desert and open ocean per unit area, though the ocean's vast area makes it roughly half of global production.
Key Insight
Change one part of an ecosystem and the effects ripple outward, as when removing wolves from Yellowstone let elk overgraze streamside willows and reshape the rivers themselves; ecosystems obey thermodynamics, since energy enters as sunlight and degrades to heat at every transfer while atoms are conserved and recycled, which is why food chains stay short and why nutrient pollution can transform a lake in a single summer.