Food Web
A food web is a network of interconnected food chains showing all the feeding relationships among organisms in an ecosystem.
Definition
A food web is a map of many food chains joined together, showing that most animals eat more than one thing and are eaten by more than one thing, such as a mouse eating seeds and insects while being hunted by hawks, foxes, and snakes, so losing that one mouse population would take a meal away from all three predators at once. More formally, a food web diagrams all the feeding relationships in an ecosystem, made of many interconnected food chains with arrows pointing from each organism to whatever eats it, and because most consumers feed on and are eaten by several species, food webs are more realistic than single chains, and webs with many connections tend to be more stable, since a consumer with several food options can switch if one source fails. At the network level, a food web represents trophic interactions as nodes for species or functional groups and directed links for energy or biomass flow, described by connectance, link density, and the distribution of trophic levels, with quantitative webs weighting links by flux; theory and experiment show that a distribution of many weak links and a few strong ones promotes stability, and that omnivory and the coupling of grazing and detrital pathways buffer ecosystems against fluctuation, while removing highly connected species or top predators can cause secondary extinctions and trophic cascades.
Example
In a North Carolina salt marsh, cordgrass and algae feed snails, fiddler crabs, and small fish, which are eaten by blue crabs, herons, and larger fish, while decomposers break down the dead cordgrass most animals cannot digest, so removing blue crabs would let snails multiply and overgraze the cordgrass. The reintroduction of wolves to Yellowstone in 1995 reduced elk browsing on willows and aspens, letting streamside vegetation, songbirds, and beavers recover, a cascade that traveled through several links of the food web, and empirical food webs typically show connectance values around $0.1$ to $0.2$.
Key Insight
A food web is really a hypothesis about how disturbance propagates, and the lesson of Yellowstone and many marine systems is that effects travel along links in ways a simple species list can never predict.